Earthquake ruptures and topography of the Chilean margin controlled by plate interface deformation
What controls the location and segmentation of mega-earthquakes in subduction zones is a long-standing problem in Earth sciences. Prediction of earthquake rupture extent mostly relies on interplate coupling models based on Global Navigation Satellite Systems providing patterns of slip deficit between tectonic plates. We here investigate if and how the strongly and weakly coupled patches revealed by these models relate to the distribution of deformation along the plate interface, i.e., basal erosion and/or underplating. From a mechanical analysis of the topography applied along the Chilean subduction zone, we show that extensive plate interface deformation takes place along most of the margin. We show that basal erosion occurs preferentially at 15 km depth while underplating does at 35 ± 10 and 60 ± 5 km depth, in agreement with P-T conditions of recovered underplated material, expected pore pressures and the spatial distribution of marine terraces and uplift rates. South of the Juan Fernández Ridge, large sediment input favors shallow accretion and underplating of subducted sediments, while along northern Chile, extensive basal erosion provides material for the underplating. We then show that, along the accretionary margin, the two last major earthquakes were limited along their down-dip end by underplating while, along the erosive margin, they were surrounded by both basal erosion and underplating. Segments with heterogeneously distributed deformation largely coincide with lateral earthquake terminations. We therefore propose that long-lived plate interface deformation promotes stress build-up and leads to earthquake nucleation. Earthquakes then propagate along fault planes shielded from this long-lived permanent deformation, and are finally stopped by segments of heterogeneously distributed deformation. Slip deficit patterns and earthquake segmentation therefore reflect the along-dip and along-strike distribution of the plate interface deformation. Topography acts as a mirror of distributed plate interface deformation and should be more systematically studied to improve the prediction of earthquake ruptures.
Predicting the extent of subduction zone seismic ruptures mostly relies on geodetic observations of interseismic coupling (e.g., Moreno et al., 2010), notwithstanding spatial resolution issues (Loveless and Meade, 2011). Some earthquakes have however ruptured low coupled zones (Noda and Lapusta, 2013) or ruptured only parts of the highly locked patches (e.g., Konca et al., 2008). This was the case for the 2014 Mw 8.4 Iquique earthquake, which only ruptured the northern end of the north Chile gap (last broken in 1877; Ruiz et al., 2014a), or the Tocopilla Mw 7.8 earthquake, which struck the down-dip and less coupled part of the same gap 7 years before (Delouis et al., 1998; Metois et al., 2016; Fig. 1). Predicting the location and extent of earthquakes clearly requires a better understanding of earthquake mechanics.
Subduction earthquake propagation has so far been either related to megathrust frictional and pore fluid pressure properties (Perfettini et al., 2010; Kaneko et al., 2010; Moreno et al., 2014), inherited stress states (Konca et al., 2008; Kaneko et al., 2010) or seafloor roughness (Kodaira et al., 2000; Wang and Bilek, 2014; Bletery et al., 2016). Megathrusts, commonly regarded as interfingered seismic and aseismic patches with contrasting frictional properties, are conventionally sub-divided into four domains with depth (A–D; Lay et al., 2012). The highly coupled domain-B hosts mega-earthquakes, while the partly coupled domain-C is characterized by moderate slip earthquakes generating significant high-frequency seismic radiation (Lay et al., 2012).
The distribution of these patches might persist over timescales of up to millions of years and control forearc morphology (Song and Simons, 2003). A spatial correlation between the coastline and the down-dip limit of the strongly locked zone has indeed been evidenced along the northern Chilean margin (Béjar-Pizarro et al., 2013; Metois et al., 2016). Coastal uplift rates inferred from marine terraces, between 0.1 and 0.3 mm a−1 over the last 400 kyr (Saillard et al., 2017), have been linked to the change of mechanical coupling between domain-B and domain-C (Saillard et al., 2017) and to underplating (Delouis et al., 1998; Adam and Reuther, 2000; Clift and Hartley, 2007). Numerical investigations recently suggested that underplating, through transient stripping of the slab-top near the base of the forearc crust, may generate periodic multi-million-year-long uplift sequences and a trackable 100 m high topographic signal (Menant et al., 2020).
Bathymetric features are considered as potential seismic barriers, based on the distribution of incoming plate roughness (Wang and Bilek, 2014; Lallemand et al., 2018) or gravimetric anomalies (Bassett and Watts, 2015). Subduction of bathymetric features induces large fracture networks, promoting removal of material from the bottom of the overriding plate (basal erosion, Ranero and von Huene, 2000) and potentially characteristic topographic features (Dominguez et al., 1998; Collot et al., 2008; Ruh et al., 2016). Heterogeneities in the structure of the damaged plate interface and stress field, as a result, would favor aseismic slip and impede the propagation of large ruptures (Wang and Bilek, 2014). Very few seismic surveys, however, have imaged seamounts surrounding a large rupture (Kodaira et al., 2000; Geersen et al., 2015).
We herein explore an alternative explanation: rather than variations of frictional properties or of seafloor roughness, earthquake segmentation might relate to the distribution of deformation along the plate interface (Wang and Bilek, 2014), as observed along strike-slip faults (Wesnousky, 2006). Both underplating and basal erosion require the redistribution of deformation, ultimately leading to the migration of the plate boundary (Vannucchi et al., 2012; Agard et al., 2018). We here show that the location of such distributed deformation along the plate interface can be captured from a simple mechanical analysis of the topography, since underplating and basal erosion both impact forearc morphology. This method is applied over a 2000 km long stretch along the Chilean margin, known to transition from erosive to accretionary from north to south, as incoming sediment thickness increases (von Huene and Ranero, 2003; Clift and Vannucchi, 2004; Clift and Hartley, 2007; Fig. 1a). This margin is one of the best instrumentally documented subduction zones, along which five mega-earthquakes, including a domain-C event, have occurred during the last 25 years (Fig. 1c). We first map out the location of the distributed deformation zones associated with basal erosion and underplating and compare them with the megathrust segmentation proposed so far. We then discuss how distributed deformation along the plate interface impacts and therefore can be used to constrain earthquake extent and mechanics.
We applied the critical taper theory (CTT; Davis et al., 1983, Fig. 2). The theory predicts the topographic slope of a wedge on the verge of failure as well as dips of its internal faults, from the frictional and pore fluid pressure properties of the wedge and megathrust (Dahlen, 1984). We used the solution of Dahlen (1984) for a non cohesive wedge, describing the critical taper, formed by the topographic slope α and the slab dip β, as a function of the angle ΨB, formed by the maximum principal stress σ1 and the base of the wedge and the angle Ψ0, formed by σ1 and the top of the wedge. The solution for the lower compressional branch is
The angles ϕint and ϕb are the internal and basal coefficients of friction defined as μint=tanϕint and μb=tanϕb, and
The internal and basal Hubbert–Rubbey fluid pressure ratios λ and λb are defined in Davis et al. (1983) as
where ρ and ρw are the wedge material and water densities and D is the water depth. The solution is exact if λ = λb and the approximation is valid for small tapers as used in this study (Wang et al., 2006).
The relationship between the topographic slope α and the slab dip β forms an envelope separating different mechanical states (Fig. 2). The wedge is at critical state if the taper formed by α and β follows a critical envelope. In that case, activation of the megathrust requires internal faulting. In contrast, if the taper exceeds this critical limit, the wedge enters in a stable domain (in the sense of the CTT), where the only possible active fault is the megathrust.
Dynamic effective friction of seismogenic zones has been shown to reach extremely low values (μ∼0.01–0.03; Fulton et al., 2013; Gao and Wang, 2014). As a consequence, a forearc above a seismogenic megathrust, with standard α and β, will systematically fall in the CTT stable domain (Fig. 2). In contrast, aseismic megathrusts are characterized by larger effective friction (μ∼0.1–0.15; Cubas et al., 2013; Gao and Wang, 2014), allowing the wedge to reach critical conditions (Cubas et al., 2013) (Fig. 2). Furthermore, at critical state, if the basal effective friction of the wedge approaches the internal effective friction, the dip of internal faults within the wedge will decrease and become parallel to the plate interface. A small difference of friction will thus lead to deformation either by basal accretion (in the lower plate) or by basal erosion (in the upper plate), instead of standard accretion characterized by thrust faults reaching the surface (Dahlen, 1984; Fig. 2). If the basal effective friction reaches the internal one, a highly fractured forearc is even expected.
We here used inversion to retrieve the effective basal friction, the internal friction and the internal pore pressure ratio from the α−β co-variation. To do so, we followed the procedure developed by Cubas et al. (2013) and detailed in the Supplement. We used the ETOPO 1 model (Amante and Eakins, 2009) to derive the bathymetric and topographic slope, and slab 2.0 (Hayes et al., 2018) for the slab dip. We built swath profiles perpendicular to the trench every 0.1∘ along trench to get average topographic slopes and slab dips with their standard deviations necessary to the inversion procedure (Fig. S1 in the Supplement). We explored from 10 to 45∘ for ϕint, from 1 to 44.9∘ for and from 0.35 to 0.975 for λ. Along the α−β profiles, we selected segments with a slope comprised between the two extreme theoretical envelops of the parameter set (Figs. 3 and S2 in the Supplement). For each of these segments, we retrieved probability density functions for the friction of the wedge, the pore pressure ratio of the wedge and the effective friction of the megathrust (Fig. S3 in the Supplement) and best misfit sets (Fig. S4 in the Supplement). We only kept segments with misfits lower than 0.1, and with values consistent with standard friction: from 25 to 43∘ for ϕint (consistent with lab experiment values (Byerlee, 1978)), from 1 to 42.9∘ for (same range as the internal friction but considering additional pore fluid pressure effect) and from 0.35 to 0.975 for λ. Figure 3a–c shows the fit between the α−β profiles and the envelops corresponding to the best misfit of the identified critical segments. These segments (Fig. 3b–d) have then been reported on Fig. 1b.
The CTT inversion only provides internal pore pressure, which can be considered as a lower bound for the megathrust pore pressure. Moreover, close effective internal and basal friction implies close internal and megathrust pore pressure.
3.1 Locating plate interface deformation
Since we seek to capture distribution of deformation along the plate interface, we compared the internal and megathrust effective friction and mapped the difference (Fig. 1b). A 1∘ difference implies a dip difference <10∘ between the megathrust and the forward-verging thrusts. This representation allows inferring areas prone to either standard accretion, or to basal erosion/accretion, the latter being herein defined as distributed deformation along the plate interface.
The transition from erosion to accretion is particularly well captured by the method despite its relative simplicity (Fig. 1a–b). North of 26∘ S (i.e., at the transition between erosive and accretionary subduction), the difference in effective friction is systematically lower than 2∘, and mostly lower than 1∘, which means that deformation is essentially located along the plate interface. The difference increases notably south of the Juan Fernández Ridge, and progresses inland towards the south, consistent with active faults mapped near the Arauco Peninsula (Fig. 1d). The method also captures basal erosion linked to the few subducted seamounts imaged around the Iquique rupture (Geersen et al., 2015; Fig. 1c).
To interpret the origin of plate interface deformation, we set the limit between accretion and interface deformation for a difference of friction angle of 1∘ and then searched for the depth distribution of each process (Fig. 4a–b). Results for different thresholds are also presented.
Accretion mostly occurs at depths shallower than 20 km, whereas distributed deformation along the plate interface is found at every depth with three favorable peaks: a first one at 15 km depth, a second at 35 ± 10 km depth and a third at 60 ± 5 km depth. Basal erosion due to bathymetric features is known to be efficient at shallow depth (von Huene et al., 2004), but might also occur slightly deeper as testified by subsidence of forearc basins (Clift and Hartley, 2007). Underplating has been observed at shallow depth (Kimura et al., 2010; Tréhu et al., 2019; Bangs et al., 2020), even for seamount moats (Clarke et al., 2018), but mostly occurs below 30–40 km, i.e., the long-term coupling–decoupling transition (Agard et al., 2018). The compilation of P-T conditions for recovered underplated material shows a similar distribution as CTT, with peaks at depths of 30 ± 5 km and 50 ± 5 km and a gap in between (Fig. 4b). Numerical modeling also suggests that erosion predominates over underplating at depths shallower than 20 km (Menant et al., 2020). As a consequence, we assume that the interface deformation documented here mostly relates to basal erosion for depths < 20 km, whereas underplating dominates at greater depths, in particular below the coast.
This depth divide between basal erosion and underplating is strengthened by estimates of the pore fluid pressure ratio (Fig. 4d–f). If low pore fluid pressure is mainly found in accretion conditions, higher fluid pressure dominates where basal erosion occurs (von Huene and Ranero, 2003; Vannucchi et al., 2012). Conversely, recovered underplated material from greater depth present very consistent P-T conditions of lithostatic pressure, which rules out any significant fluid overpressure (Agard et al., 2018). The first peak for basal erosion at low pore fluid pressures in Fig. 4e could be related to the 20 km depth assumption and reveal the minor occurrence of underplating at shallower depth. The trimodal distribution deduced from CTT (one peak for basal erosion and two peaks for underplating) and differences in pore pressure remains for a 0.5∘ or a 2∘ difference of friction.
Most of the critical segments attesting to distributed interface deformation have lengths smaller than 7 km, and do not exceed 20 km (Fig. 4c). This is in good agreement with the length of graben and horst and seamounts observed on the subducting plate (von Huene and Ranero, 2003; Geersen et al., 2015). This is also consistent with seismic observations (Kimura et al., 2010; Tréhu et al., 2019; Bangs et al., 2020) and the long-term rock record showing generally ≤ 300–500 m thick exhumed slices from the top of the slab (Agard et al., 2018) extending 5–10 km downdip at most (Plunder et al., 2012, 2013). Such tabular shape ratios, with seamounts probably near the higher bound (Bonnet et al., 2019), further support interface deformation parallel to the megathrust (Fagereng and Sibson, 2010; Rowe et al., 2013).
3.2 Relating plate interface deformation with long-term coastal uplift
In order to approach internal effective friction, the basal friction of the wedge needs to be relatively high (Fig. 2a). Deformation captured at depth by CTT therefore corresponds to underplating in the making, when deformation is still distributed: once tectonic slicing is achieved, basal friction will reach low values characteristic of mature faults, bringing the taper in the stable domain and impeding detection by CTT.
We here show that active, incipient underplating takes place along most of the Chilean margin (Figs. 1, 5a), whether or not strain accumulation ultimately turns these critical zones into tectonic slices (Agard et al., 2018; Bonnet et al., 2019; Kimura et al., 2007). Along the Tocopilla and Maule segments, incipient underplating takes place below the slab – continental Moho intercept (orange curve on Fig. 1c), whereas it is located above and near the intercept between 25 and 33∘ S. Along the northern erosive margin (Fig. 5a–b), the thin oceanic sediment is covered together with eroded material, and pieces stripped from the subducting plate are being possibly underplated. Southward, larger sediment input favors shallow accretion and underplating of subducted sediments (Fig. 5a–b), as locally observed by seismic surveys (Tréhu et al., 2019).
To evaluate the possible link between coastal uplift and regions of distributed deformation along the plate interface, we compare our spatial distribution of underplating with documented marine terraces and uplift rates (Saillard et al., 2017; Fig. 5a–c). While the irregular distribution of marine terraces may relate to fragmentary preservation rather than discontinuous underplating, most of them coincide with segments of underplating (or with standard accretion near the Arauco peninsula). The typical length of the segments showing distributed deformation along the interface is modest (∼ 10–20 km). This advocates for propagation of deformation only into the topmost part of the subducting plate, e.g., into the stack of oceanic/eroded material or along a hydrated basalt layer within the crust. Although the comparison with uplift rates inferred from morphometric analysis and modeling of landscape evolution (Melnick, 2016) shows no particular pattern, underplating of relatively thin slices would be consistent with both the low uplift rates and the rock record (Agard et al., 2018).
Shallow normal faults identified along northern Chile have been related to potential underplating (Adam and Reuther, 2000; Clift and Hartley, 2007). We here show that they are located above underplating areas, and that normal faults transition southward to thrusts where accretion prevails (Fig. 5a–c).
3.3 Large megathrust earthquakes surrounded by plate interface deformation
We now compare zones of distributed deformation with areas struck by recent large earthquakes (Fig. 1c). The Mw 8.1 2014 Iquique earthquake is surrounded by patches of distributed deformation associated with basal erosion according to their depth and to seamounts detected in the area (Geersen et al., 2015). The aftershock, located 50 km south of the main event, also falls in between patches of distributed deformation. The Mw 7.8 2007 Tocopilla earthquake is limited up-dip as well as to the south by interface deformation attributed to underplating. The Mw 8.1 1995 Antofagasta earthquake seems limited up- and down-dip by distributed deformation along the plate interface. The high slip areas of the Mw 8.2 2015 Illapel and the Mw 8.7 2010 Maule earthquakes are both limited down-dip by plate interface deformation. Their frontal extent partially propagates in areas at critical state but corresponding to standard accretion. A small portion of the earthquake ruptures do overlap with the plate interface deformation, but high slip areas are always clearly limited by plate interface deformation. Moreover, Fig. 6 shows the diversity of published co-seismic models, emphasizing the uncertainties related to the proposed slip areas, that need to be taken into account for the comparison.
To summarize, along the accretionary part of the margin, mega-earthquakes are delimited down-dip by underplating whereas along the erosive domain, the extent is controlled by both basal erosion and underplating.
We now investigate the relationships between the critical segments characterized by relatively high effective basal friction, hence expected to behave aseismically, both with the along-strike segmentation inferred for large historical earthquakes (Mw ≥ 7.5; Saillard et al., 2017, Fig. 5a–d) and with the estimates of coupling (Metois et al., 2016; Klein et al., 2018, Fig. 5a–e). Segments with limited plate interface deformation (< 15 km) coincide with a significant number of rupture terminations (blue and gray overlays, respectively: Fig. 5a–d), particularly in the southern region. Between 29 and 24∘ S, rupture terminations coincide with segments where plate interface deformation is reduced. The underplating segments also show a long wavelength distribution (4 to 8∘ latitude) corresponding to the three main seismic regions visible on Fig. 5a, d (respectively, located north of Mejillones, between Mejillones and Punta Choros, and to the south of Punta Choros). No correlation exists with sediment input (Fig. 5b). The proportion of a swath profile at critical state is then compared to the average estimate of coupling along this profile (Fig. 5e). A positive correlation is evidenced at small scale (1.5 and 2∘ latitude, Fig. S7 in the Supplement) between the segments with minimal underplating (blue overlays; Fig. 5a) and the regions of minimal coupling (red overlays; Fig. 5e). On a larger scale (4 and 8∘ latitude, Fig. S7), an anti-correlation is observed with coupling: zones of high coupling correspond to domains where underplating is somewhat more limited (e.g., between 31 and 38∘ S). As shown by Fig. 1d, areas marked by intense plate deformation are mostly located down-dip of the highly locked patches, in particular in the south. Continuous and voluminous sediment influx provides spatially and temporally stable mechanical conditions (Olsen et al., 2020). This leads to stationary seismic asperities followed, once the Moho is crossed, by the regular underplating of very thin and relatively short slices (typically 300–500 m thick, 5–10 km long). This spatial and temporal stability promotes topographic build-up, explaining the spatial coincidence between strongly coupled patches and efficient underplating. In contrast, segments with limited underplating might reveal spatially and temporally heterogeneous conditions along the megathrust (Olsen et al., 2020), i.e., heterogeneous deformation impeding earthquake propagation and significant topographic build-up.
Noteworthily, the hypocenters of the last major events lie at the boundary with a patch of distributed deformation. Finally, we show that the high-frequency radiation generated by the Maule and Illapel earthquakes (Wang and Mori, 2011; Meng et al., 2015, 2018) coincide with areas of extensive interplate deformation (Fig. 1c), along which spatially restricted fault planes might have been dynamically triggered.
4.1 Link between earthquake ruptures and plate interface deformation
We herein show two major results: (1) recent earthquakes are bounded by extensive plate interface deformation characterized, along the southern accretionary part of the margin by underplating at their down-dip edge and, along the northern erosive part of the margin by both basal erosion and underplating (Fig. 1); (2) along strike, segments characterized by minimal interplate deformation largely coincide with lateral earthquake terminations (circles; Fig. 5d). These observations demonstrate a close link between distributed interplate deformation and earthquake segmentation.
We therefore propose that plastic deformation and stress build-up associated with interplate deformation along distributed fault planes of limited extent (Fig. 7 – step 1) eventually lead to mega-earthquake nucleation (Fig. 7 – step 2). This is suggested by the location of hypocenters at boundaries with distributed deformation (Fig. 1). This interseismic deformation might be accompanied by some micro-seismicity, as observed prior to the Iquique rupture (Schurr et al., 2020) and in the upper plate in northern Chile (Sippl et al., 2018). If high pore fluid pressure prevails, this deformation could also be accompanied by transient seismic/slip events (Liu and Rice, 2007; Kimura et al., 2010; Collot et al., 2017), which agrees with observations for the Iquique earthquake (Ruiz et al., 2014a; Meng et al., 2015; Socquet et al., 2017) and others elsewhere (Tohoku, Kato et al., 2012; Guerrero, Radiguet et al., 2016).
Once nucleated, large earthquakes propagate along well-localized and smoothed rate-weakening fault planes (Bletery et al., 2016) limited by elongate zones of underplating which, in addition, inhibit further rupture propagation and slip (Figs. 1, 7 – step 3).
These earthquakes abut regions of heterogeneously distributed deformation and stress concentrations (Fig. 7 – step 4), impeding the development of well-localized slip zones (Wang and Bilek, 2014), and again significant slip (Wesnousky, 2006). Some of these fractures might however slip seismically during large events and produce the observed high-frequency radiation (Meng et al., 2018, 2015).
4.2 Down-dip segmentation and plate interface deformation
Within this framework, domain-C (Lay et al., 2012) would correspond to the region of incipient underplating, and domain-B to the smoothed rate-weakening fault zone. This is consistent with partial coupling of domain-C, where deformation is dominated by creep mechanisms, and elastic strain is accumulated along undeformed segments of the megathrust, producing moderate slip earthquakes with a higher recurrence time and significant coherent short-period seismic radiation. Domain-C might more specifically be delimited by the two underplating peaks (Fig. 4), the upper one corresponding to the slicing of oceanic sediments and/or eroded continental material, the second to the propagation of deformation into the hydrated basalt layer once the continental Moho is crossed (Agard et al., 2018), a process ultimately leading to the underplating of very thin and relatively short slices (typically 300–500 m thick, 5–10 km long; Agard et al., 2018).
Data suggest that domain-C, by locally concentrating interplate permanent deformation due to specific geometry (i.e., morphological asperities) and/or mechanical behavior (i.e., frictional properties, fluid content, porosity), shields the rate-weakening domain-B from long-lived deformation (Fig. 7). Contrary to the prevalent paradigm, domain-B should not be regarded as an “asperity”, i.e., a strongly coupled yet mechanically weak zone with high recurrence time earthquakes, but instead as a relatively weak zone devoid of permanent deformation which only stores elastic energy. In this new paradigm, loading along domain-C tightly controls slip deficit as well as rupture nucleation and extent along domain-B (Fig. 7), explaining the correlation observed between historical earthquakes, earthquake terminations and distributed plate interface deformation (Fig. 5a, d). In the short term, elastic deformation generates subsidence at the coast during domain-B earthquakes and uplift for domain-C events (Melnick, 2016). In the long term, the thin underplated slices ultimately build a topographic signal over millions of years of slow uplift, particularly difficult to capture with geodetic measurements due to the interplay of elastic and plastic deformation (Menant et al., 2020). The Coastal Cordillera present along much of the Chilean margin could result from this long-term underplating process. The absence of upper plate deformation above the seismogenic zone in conjunction with topographic build-up above underplating areas account for the correlation between seismic behavior and gravity anomalies (Song and Simons, 2003).
We herein show, through a simple CTT analysis of topography, that, along the Chilean subduction zone, earthquakes are bounded by long-lived, extensive plate interface deformation occurring preferentially at 35 ± 10 and 60 ± 5 km depth. Along the central Chilean margin, earthquakes are limited along their down-dip edge by underplating while, along northern Chile, they are surrounded by distributed deformation related to both basal erosion and underplating. Plate interface deformation might control earthquake nucleation, extent and arrest, and its distribution provides a relatively reliable image of the seismic potential at depth. This topography analysis may also remedy the lack of preserved marine terraces or weakly constrained coupling models. This method could therefore be used more systematically to identify regions with precursory transient signals and to enhance predictions of future earthquakes' extent.
Data are provided in the Supplement. The code is available on demand.
See the Supplement for details on the method, supporting figures, and data. The supplement related to this article is available online at: https://doi.org/10.5194/se-13-779-2022-supplement.
NC designed the study; NC and RT ran the simulations; NC, RT, PA analyzed the results; NC and PA wrote the paper.
The contact author has declared that neither they nor their co-authors have any competing interests.
Publisher’s note: Copernicus Publications remains neutral with regard to jurisdictional claims in published maps and institutional affiliations.
We thank the two anonymous reviewers for their helpful comments.
This research has been supported by the Agence Nationale de la Recherche (grant no. ANR-13-PDOC-0013–01).
This paper was edited by Stefano Tavani and reviewed by two anonymous referees.
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